As follows from our interpretation, the appearance of a huge voltage gradient (evoked by the sudden rush-in current) represents the output information the neuron delivers, and also the input information it receives from its upstream neurons through its synapses. Given that the input currents delivered by the spikes are gated by the neuron, the information that can be accounted in the computation must be in the front side of spikes. (The back side is mainly needed for restoring the resting potential.)
The front side (in the form of a sudden step in the value of the voltage gradient) delivers an extremely precise timing information about at what time the rush-in event in the sender happened, explaining the half-understood experience [143] why ”the timing of spikes is important with a precision roughly two orders of magnitude greater than the temporal dynamics of the stimulus”. If exceeding the threshold is the consequence of the charge arriving from a single upstream neuron, the neuron simply transmits the timing information it received. If several smaller gradients are summed (and recall that the component gradients decay with time after their arrival) for reaching the gradient, then their information content is weighted.